Look man according to the wording of the question, the wheels would end up spinning infinitely fast before the plane left the ground. In reality it is not possible to build a treadmill to take off from, but in theory, the plan would take off anyway since it is not thrusting against the ground, but the air, which is not affected by the treadmill.
As Kev so aptly stated:
As Kev so aptly stated:
IF THE TREADMILL WAS GOING A MILLION TRILLION BAJILLION GAGILLION MILES PER NANOSECOND IN THE OPPOSITE DIRECTION THE PLANE WOULD STILL FVCKING TAKE OFF.