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Upgrade from a R290.....

980 Ti on the Nvidia side pretty much.
 
Or just wait till next year as I'm sure buyers remorse will kick in. 980ti is a good choice but I'm sure pascal will offer improved features and performance at a lower cost and even more performance for the successor to said card.
 
From my sig below, you can see I own BOTH a GTX980TI (EVGA GTX980TI SC) and 2 Sapphire Tri-X R9 290s. If you want to stay away from AMD, the GTX980TI seems to be the only "upgrade".
 
A B stock gtx980 with overclock for 350$ is a good upgrade.
http://www.evga.com/Products/Product.aspx?pn=04G-P4-1980-RX

The overclock should net you about 18% more performance with games.
http://www.anandtech.com/show/8526/nvidia-geforce-gtx-980-review/22

That's a 37%+ upgrade.
perfrel_1920.gif
 
Do you currently have a reference 290, or is it an aftermarket model?
What's your reason for wanting the upgrade? If it's just more FPS, going to a 980 likely isn't going to give you a worthwhile increase, especially if you have an aftermarket cooler and can overclock it.
If it's that you want to get rid of a loud reference blower and get a little more performance, or just really hate CCC, the B-stock 980 might be an option. Going rate on a 290 seems to be around $175, so it will cost you ~180-220 (depending on shipping, taxes and whatnot) to get a 20% performance increase. Unless you really hate running that 290 you'd be better off just buying another 290 and running them in xfire until Pascal arrives.

Going with a 980 Ti would set you back 425-475 out of pocket, but you'd probably see a 60-80% performance increase or so depending on overclocking, which would be much more noticeable.
 
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