Originally posted by: 0marTheZealot
Originally posted by: PJABBER
Originally posted by: 0marTheZealot
Originally posted by: PJABBER
Based on my review of the technical studies, I do believe CO2 to be of minimal environmental impact, unless it is maybe of positive benefit in supporting the growth of plant life, the basis of all life on this planet. Maybe methane and NO have more of an injurious impact, in my reading the jury is still out on those as well.
You should learn some plant physiology. Look up photorespiration, it's a phenomena in which plants actually respire (that is release CO2) due to high levels of CO2. The main photosynthesis enzyme, Rubisco, is inhibited at higher CO2 concentrations. IIRC, around 400ppm CO2, photosynthesis in a large number of plants completely shuts down.
Those are laboratory conditions and far, far, far from existing and forecast levels even under the absolute worst case scenarios.
Trends in Atmospheric Carbon Dioxide
For example, the last three years measurements of CO2 PPM -
Mauna Loa, Hawaii
2006 1.69
2007 2.17
2008 1.66
Sea Level - Global Mean
2006 1.77
2007 2.12
2008 1.79
Do you even analyze the data you just linked me? Staring at me, right in my face, is the measurements for CO2 that says in 2008, Mauna Loa had a mean CO2 level, in PPM, of ~380. Not 1.66. That is the growth rate with respect to ppm/year. That is, on average, you can expect the amount of CO2 to go up by 1.66ppm year on year measurement.
You are right! I was distracted by a phone call reminding me that I have an important dinner engagement this evening. Multi-tasking fails me again!
Let me see if I can get a handle on your fears -
Regulation by carbon dioxide. Since carbon dioxide and oxygen compete at the active site of RuBisCO, carbon fixation by RuBisCO can be enhanced by
increasing the carbon dioxide level in the compartment containing RuBisCO (chloroplast stroma). Several times during the evolution of plants, mechanisms have evolved for increasing the level of carbon dioxide in the stroma (see C4 carbon fixation). The use of oxygen as a substrate is an apparently-puzzling process, since it seems to throw away captured energy. However it may be a mechanism for preventing overload during periods of high light flux. This weakness in the enzyme is the cause of photorespiration, such that healthy leaves in bright light may have zero net carbon fixation when the ratio of O2 to CO2 reaches a threshold at which oxygen is fixed instead of carbon. This phenomenon is primarily temperature-dependent. High temperature decreases the concentration of CO2 dissolved in the moisture in the leaf tissues. This phenomenon is also related to water stress. Since plant leaves are evaporatively cooled, limited water causes high leaf temperatures. C4 plants use the enzyme PEP carboxylase initially, which has a higher affinity for CO2. The process first makes a 4-carbon intermediate compound, which is shuttled into a site of C3 photosynthesis then de-carboxylated, releasing CO2 to boost the concentration of CO2, hence the name C4 plants.
Role of photorespiration
Photorespiration is said to be an evolutionary relic. Photorespiration lowers the efficiency of photosynthesis by removing carbon molecules from the Calvin Cycle. The early atmosphere in which primitive plants originated contained very little oxygen, so it is hypothesized that the early evolution of RuBisCO was not influenced by its lack of discrimination between O2 and carbon dioxide. Although the functions of photorespiration remain controversial, it is widely accepted that this pathway influences a wide range of processes from bioenergetics, photosystem II function, and carbon metabolism to nitrogen assimilation and respiration. Crucially, the photorespiratory pathway is a major source of H2O2 in photosynthetic cells. Through H2O2 production and pyridine nucleotide interactions, photorespiration makes a key contribution to cellular redox homeostasis. In so doing, it influences multiple signaling pathways, particularly those that govern plant hormonal responses controlling growth, environmental and defense responses, and programmed cell death.
Another theory postulates that it may function as a "safety valve", preventing excess NADPH and ATP from reacting with oxygen and producing free radicals, as these can damage the metabolic functions of the cell by subsequent reactions with lipids or metabolites of alternate pathways.
Since photorespiration requires additional energy from the light reactions of photosynthesis, some plants have mechanisms to reduce uptake of molecular oxygen by RuBisCO. They increase the concentration of CO2 in the leaves so that Rubisco is less likely to produce glycolate through reaction with O2.
C4 plants capture carbon dioxide in cells of their mesophyll (using an enzyme called PEP carboxylase), and oxaloacetate is formed. This oxaloacetate is then converted to malate and is released into the bundle sheath cells (site of carbon dioxide fixation by RuBisCO) where oxygen concentration is low to avoid photorespiration. Here Carbon dioxide is removed from the malate and combined with RuBP in the usual way. The Calvin cycle then proceeds as normal.
The enzyme PEP carboxylase (which catalyzes the combination of carbon dioxide with a compound called Phosphoenolpyruvate or PEP) is also found in other plants such as cacti and succulents who use a mechanism called Crassulacean acid metabolism or CAM in which PEP carboxylase sequesters carbon at night and releases it to the photosynthesizing cells during the day. This provides a mechanism for reducing high rates of water loss (transpiration) by stomata during the day.
This ability to avoid photorespiration makes these plants more hardy than other plants in dry and hot environments where stomata are closed and internal carbon dioxide levels are low. C4 plants include sugar cane, corn (maize), and sorghum.
OK, now it seems that high Co2 is actually a good control/balance for differing levels of oxygen.
Despite previous reports of no apparent photorespiration in C4 plants based on measurements of gas exchange under 2 versus 21% O2 at varying [CO2], photosynthesis in maize (Zea mays) shows a dual response to varying [O2]. The maximum rate of photosynthesis in maize is dependent on O2 (approximately 10%). This O2 dependence is not related to stomatal conductance, because measurements were made at constant intercellular CO2 concentration (Ci); it may be linked to respiration or pseudocyclic electron flow. At a given Ci, increasing [O2] above 10% inhibits both the rate of photosynthesis, measured under high light, and the maximum quantum yield, measured under limiting light ([phi]CO2). The dual effect of O2 is masked if measurements are made under only 2 versus 21% O2. The inhibition of both photosynthesis and [phi]CO2 by O2 (measured above 10% O2) with decreasing Ci increases in a very similar manner, characteristically of O2 inhibition due to photorespiration. There is a sharp increase in O2 inhibition when the Ci decreases below 50 [mu]bar of CO2. Also, increasing temperature, which favors photorespiration, causes a decrease in [phi]CO2 under limiting CO2 and 40% O2. By comparing the degree of inhibition of photosynthesis in maize with that in the C3 species wheat (Triticum aestivum) at varying Ci, the effectiveness of C4 photosynthesis in concentrating CO2 in the leaf was evaluated. Under high light, 30[deg]C, and atmospheric levels of CO2 (340 [mu]bar), where there is little inhibition of photosynthesis in maize by O2, the estimated level of CO2 around ribulose-1,5-bisphosphate carboxylase/oxygenase (Rubisco) in the bundle sheath compartment was 900 [mu]bar, which is about 3 times higher than the value around Rubisco in mesophyll cells of wheat. A high [CO2] is maintained in the bundle sheath compartment in maize until Ci decreases below approximately 100 [mu]bar. The results from these gas exchange measurements indicate that photorespiration occurs in maize but that the rate is low unless the intercellular [CO2] is severely limited by stress.
Light-dependent carbon dioxide release and oxygen uptake in photosynthetic organisms caused by the fixation of oxygen instead of carbon dioxide during photosynthesis. This oxygenation reaction forms phosphoglycolate, which represents carbon lost from the photosynthetic pathway. Phosphoglycolate also inhibits photosynthesis if it is allowed to accumulate in the plant. The reactions of photorespiration break down phosphoglycolate and recover 75% of the carbon to the photosynthetic reaction sequence. The remaining 25% of the carbon is released as carbon dioxide. Photorespiration reduces the rate of photosynthesis in plants in three ways: carbon dioxide is released; energy is diverted from photosynthetic reactions to photorespiratory reactions; and competition between oxygen and carbon dioxide reduces the efficiency of the important photosynthetic enzyme ribulose-bisphosphate (RuBP) carboxylase. There is no known function of the oxygenation reaction; most scientists believe it is an unavoidable side reaction of photosynthesis.
The rate of photosynthesis can be stimulated as much as 50% by reducing photorespiration. Since photosynthesis provides the material necessary for plant growth, photorespiration inhibits plant growth by reducing the net rate of carbon dioxide assimilation (photosynthesis). Plants grow faster and larger under nonphotorespiratory conditions, in either low oxygen or high carbon dioxide atmospheres. Most of the beneficial effects on plant growth achieved by increasing CO2 may result from the reduced rate of photorespiration.
There are some plants that avoid photorespiration under certain conditions by actively accumulating carbon dioxide inside the cells that have ribulose-bisphosphate carboxylase/oxygenase. Many cacti do this by taking up carbon dioxide at night and then releasing it during the day to allow normal photosynthesis. These plants are said to have crassulacean acid metabolism (CAM). Another group of plants, including corn (Zea mays), take up carbon dioxide by a special accumulating mechanism in one part of the leaf, then transport it to another part of the leaf for release and fixation by normal photosynthesis. The compound used to transport the carbon dioxide has four carbon atoms, and so these plants are called C4 plants. Plants that have no mechanism for accumulating carbon dioxide produce the three-carbon compound phosphoglycerate directly and are therefore called C3 plants. Most species of plants are C3 plants.
So low C02 is bad and high CO2 is actually good. Right?